The ruthless queen of the Pure Ones who enslaves young owls as soldiers and metal pickers. She orders the brainwashing process, executes dissenters like Grimble, commands the final assault, and retreats after Metalbeak's death, with hints she remains a threat.
Natural leader of the Guardians opposed to the Pure One queen.
Kludd pledges allegiance to Nyra, seeking validation through military service.
Historical enemies connected through past battles and lost loved ones.