While working as a part-time security guard for the Kougami Foundation's art museum, nomadic vagabond Eiji Hino is knocked out by two thieves. As they rob the artifact chamber, a mass of coins forms into a disembodied right arm and breaks the seal on a sarcophagus, releasing coin-like monsters called the Greeed. Upon being alerted to the robbery, the foundation sends private security to trap the Greeed, but the latter kill all except for Shintaro Goto while escaping. Sometime later, Eiji is awok
As Kougami Foundation robots collect the Kamakiri Yummy's remains for the organization's leader, Kousei Kougami, Ankh tells Eiji he is keeping Shingo alive and reveals that he requires "Cell Medals", like the ones the Yummy was made of, to rebuild his body and Core Medals, which give him and the Greeed life and power. Meanwhile, Uva finds the art thieves and uses one of them to create a new, blank Yummy that hungers for wealth. Sensing the monster's creation, Ankh leads Eiji to the bank, where t
As Ankh reviews Shingo's memories to determine who Hina is, she notices the former's arm and tears it off with her superhuman strength, but faints in shock. Ankh renews his attempt to kill her until Eiji threatens to throw his Rider equipment away. Ankh reluctantly changes his mind before Eiji takes him to the Cous Coussier restaurant, but the former leaves. Meanwhile, one of the Greeed's members, Kazari, notices how much humanity has changed since they were sealed and uses the gluttonous Monta
As the Neko Yummy escapes, Kazari forces Eiji not to pursue and offers to make a deal with Ankh, who asks for time to consider it. As Eiji collapses from exhaustion, Hina brings him to Cous Coussier, where he reveals everything he has become involved in to her and promises to save Shingo while recovering. Using the internet, Ankh locates Fukuji. After learning from Shiraishi of Fukuji's presence at a nearby restaurant, a still injured Eiji leaves to stop him despite Hina's pleas. As he, Ankh, an
Kougami negotiates with Eiji and Ankh, the latter especially, to form a partnership wherein Kougami receives 70% of any Cell Medals acquired. Though Ankh refuses, Kougami gives him time to consider. Meanwhile, the Greeed assume human forms based on Kazari's findings while one of their number, Mezool, uses a shopaholic and classmate of Hina's named Haruka Yamano to create a hidden mass of roe that gradually multiply based on her excessive shopping. The next day, Ankh attempts to conduct research
A weakened Uva reluctantly withdraws, vowing revenge on Ankh and his fellow Greeed. Ankh reveals to Eiji that the Greeed, at their strongest, possess three sets of unique Core Medals and if Eiji were to use one set, he can emulate their full power. Ankh leaves Eiji with enough medals to use his regular Rider forms before leaving to locate Kougami. During their subsequent meeting, Kougami agrees to receive 60% of all medals acquired, which Ankh begrudgingly agrees to. Meanwhile, Uva confronts Kaz
After establishing a hideout, Kazari tells Mezool that he intends to use their fellow Greeed, Gamel, to regain their stolen Core Medals while Uva enacts a separate plan to destroy Ankh and an amused Gamel becomes inspired by a bickering married couple to create the Bison Yummy to satiate his desire to see people being hit with objects. Concurrently, Eiji takes up part-time work at Cous Coussier despite Shiraishi offering him full-time and housing before a woman named Momoko Komori arrives, looki
Amidst Hino's failed attempts at stopping the Komoris' argument, Momoko takes the winning scratchcard and throws it into a nearby river. As Takeshi frantically searches for it, she reveals to Hino why Takeshi has become so greedy. In time, Hino helps the Komoris reconcile, with Takeshi resuming his photography career. Meanwhile, a weakened Ankh escapes, but finds he is unable to heal. Refusing to call Hino out of pride, he waits for Shingo's body to become available. Concurrently, Mezool stops U
In flashbacks, Eiji fails to save a girl amidst a foreign civil war. In the present, Satonaka meets with Eiji to collect Kougami's fee until an explosion occurs outside. While investigating, Eiji discovers a strange man with a puppet on his shoulder and a Same Yummy, which caused the explosion, though both escape. To his dismay, Eiji discovers it is connected to more Same Yummy that were created by Mezool from someone who desires destruction. Meanwhile, as they feast on the Same Yummy, Kazari de
As Maki observes the bomber, Kougami Biotech Laboratory technician Toru Tadano, Ankh scolds Eiji, who later discovers Maki's whereabouts. The pair travel to the laboratory, where Ankh identifies Tadano as the Same Yummies' host while Eiji destroys another instance of the Yummy. Spotting Eiji, Maki puts the laboratory on lockdown to prevent him from interfering with the nest. While Eiji leaves to stop Tadano, Goto locates the nest and fights off the emerging Same Yummies. Upon defusing the bombs
Eiji encounters an old friend and globetrotting blogger, Keisuke Tsukuba, who has come to Japan to find a publisher for his work. Eiji offers to help, but Tsukuba is rejected in favor of a better writer named Kotaro Sawada. Taking advantage of this, Uva finds Tsukuba and uses him to create the Ageha Yummy, which siphons Sawada's talent and transfers it to Tsukuba. Upon learning of the attack, Eiji fights the Yummy, but it escapes. Having seen this, an upset Goto reports Eiji's failure to Kougami
Following Goto's attack, the Ageha Yummy retreats again while Eiji learns of Goto's distaste towards him. Just then, having successfully evaded Uva, Hina arrives to tell Eiji of what happened to Tsukuba, who has become a minor celebrity. As Ankh considers stealing Gamel's Core Medals, Eiji and Hina confirm Tsukuba is the Ageha Yummy's host after Tsukuba reveals he seeks to prove himself and asks to use Eiji's story as his own. After learning Gamel recovered two of Mezool's Core Medals, Uva offer
Satonaka meets with Goto to inform him that he will be replacing her as Kougami's aide while she takes a vacation. Concurrently, Ankh struggles with not being able to find his personal Core Medals while Kazari uses Kei Tamura, a surgeon who feels that her talents are being wasted, to create a new Yummy, causing her to perform unauthorized operations on her own. Sensing the Yummy's creation, Ankh uses Shingo's memories and Eiji and Hina's assistance to infiltrate the hospital Tamura works at by p
As the hospital staff take Shingo into their care, the director suffers a heart attack upon learning what happened to Tamura and reveals to Eiji that he held her back because of her pride and lack of fear for her patients. Resolving to save Tamura, Eiji leaves to find her and the Siamneko Yummy, successfully doing so before it can kill Ankh. Eiji frees Tamura and convinces her to reconcile with her father while Ankh returns to Shingo's body and Tamura operates on her father. Though Kazari interv
Ankh reveals to Eiji that he has been spying on Kougami and learned Goto is going to oversee a truck containing 5000 Cell Medals and one of Ankh's Core Medals. Eiji refuses to help until Kazari arrives, revealing further that Hina was placed in the truck. Upon confirming her disappearance, Eiji convinces Goto to let him follow the truck. However, they are attacked by the other Greeed, who intend to steal the Cell Medals. Ankh arrives to help fight them and get his Core Medal back, but Eiji loses
Centuries prior, alchemists created the Greeed, who were born with an insatiable desire to become whole. In the present, having formed an alliance with Maki, who wants to see what happens when a Greeed absorbs every Core Medal, Kazari tricks Gamel into absorbing Mezool's, restoring his full power. Maki asks for Kazari's medals, but he refuses, taking Ankh's medal as payment. Concurrently, Ankh and Goto separately confront Kougami and Maki, with the former revealing the latter's role in what happ
Birth introduces himself as Akira Date and, using the Birth system instruction manual, identifies Eiji and Ankh before leaving. Ankh grows irate over not being able to research Date, but Eiji assures him that they will learn more about him soon enough. Meanwhile, a girl named Rie Shiratori's desire to perform better for her kendo class attracts Uva, who uses her to create the Kabuto Yummy, which attacks everyone who is better than her at kendo. Upon learning of it, Eiji tries to fight the monste
Upon being released from the hospital, Rie returns to her kendo class to see her teacher, Masaru, but is stopped by Uva, who tells her to wait for her desire to come true. Meanwhile, as they discuss the Kuwagata Yummy's creation, Ankh and Eiji realize the Kabuto Yummy actually represented her desire to reach the goal that the Kuwagata Yummy represents. Concurrently, Goto formally introduces himself to Date to learn more about the Birth system, recalling how Kougami did not let him use it because
Three years prior, a criminal named Toichi Yamagane was betrayed by his partner Yasuji "Yasu" Okumura and arrested by Shingo, for which he vowed revenge on them both. In the present, Eiji accidentally sleeps in and arrives late to Cous Coussier, but finds Goto is working there now. Later, while attending a meeting with Kougami, Eiji receives one of Ankh's Core Medals and is told that he needs to become stronger as the remaining Greeed are too. Afraid that Ankh will take his medal and leave Shing
Kazari takes the last medal he needs from Eiji, but Ankh anticipated his plan, worked with Yasu to facilitate it, and offers to trade one of his own medals for Kazari's last medal before claiming that Eiji has it and throwing him off the roof of a building. As Kazari and the Lion-Kurage Yummy pursue him, Hina takes Ankh's medal to force him to help Eiji. He threatens to kill her, but ultimately cannot follow through. Concurrently, Date trains Goto in using the Birth system while Yasu saves Eiji
While helping the Cous Coussier staff prepare for Valentine's Day, Goto witnesses an older man harassing a young boy, Takeshi, over the latter telling him to stop littering. Soon after, Takeshi's father, Susumu Kanbayashi, arrives to congratulate his son. However, Takeshi runs off after Susumu refuses to come home to him and his mother. Uva subsequently attacks Susumu, using his desire for justice to create the Batta Yummy. Meanwhile, Eiji takes Ankh on an errand to help him take his mind off no
Susumu tells Eiji to stop interfering before he, Takeshi, and the Batta Yummy flee to attack previously untouchable criminals, with Uva watching over them. Later, while continuing his training with the Birth system, Goto recalls Takeshi's words and realizes justice can have a corrupting influence if taken to an obsessive degree. He later asks why Date fights, but the latter instead advises him to believe in himself before leaving to fight the Batta Yummy. He joins Eiji in confronting the monster
After absorbing his share of Gamel's medals, Kazari secretly uses a Beauty Marine Lab beauty salon employee named Yumi Sakura to create a Yummy egg and meets with Uva to taunt him over not absorbing his share of Mezool and Gamel's medals. A fight ensues, though Uva escapes. Meanwhile, as Ankh grows worried about the state of the Core Medals and Eiji not having enough power, Maki advises Date to undergo a check-up for potential side effects of using the Birth system. Date meets with Eiji to discu
With Eiji lovesick over Yumi's otherworldly beauty, the unaffected Date and Ankh try to find the Yummy egg, only to be thwarted by the Ei Yummy. The next day, Goto helps Hina and Ankh bring Eiji back to his senses and take one of Mezool's medals from Kazari while Date meets with Kougami to discuss his health, current Cell Medal total, and an upgrade to the Birth system that Maki is developing. As the Ei-Sai Yummy emerges from the egg, it joins Yumi in capturing Rei so she can take her place as M
Eiji attempts to ask Ankh about his nature as a Greeed, but the latter storms off. Meanwhile, the Cous Coussier staff play host to a party led by Kazuki Okamura, a punch-drunk boxer who is being forced to retire. Date, Goto, and Eiji later learn of an Oumu Yummy healing Okamura's arm by petrifying that of his friend's. With Ankh watching from afar, Eiji and Date transform to fight the monster until it flies off with Okamura. After sending a Kougami Foundation robot to track it, Goto interrogates
As Date tends to his injuries, Eiji remains convinced that a Greeed with Ankh's medals created the Oumu Yummy despite what happened and realizes that the Yummy steals attributes from others and gives it to their host. Date and Goto leave Eiji to recover while they look for the Yummy, but Eiji joins them anyway. The trio eventually find a fully healed Okamura and the Oumu Yummy attacking Ankh in an attempt to steal his medals. Joining the fray, they destroy the Yummy and undo its effects on Okamu
Kougami summons Eiji and Ankh to help him celebrate the 1000th episode of the Kamen Rider franchise by making an anniversary film, Kamen Rider OOO vs Shocker, with Date serving as director. In particular, he promises to tell Ankh who has the rest of his medals. As Date fills out the cast, Ankh realizes Kougami Foundation employee In Sendo, who has been cast as a Shocker foot soldier, is a Yummy host. Before he can act on this, he detects another Oumu Yummy and has Eiji and Date join him in stopp
Despite destroying the Oumu Yummy, Eiji and Date are overpowered by the Ika-Jaguar Yummy before it escapes. Meanwhile, Ankh confronts Kougami for information on who has his Core Medals, only to later learn Kougami lied. Despite his injuries, an undeterred Date continues production on the film. However, Sendo leads an army of monsters in attacking the production team, revealing himself as an actual, formerly retired Shocker foot soldier. Though Eiji destroys the monsters, Kazari and the Ika-Jagua
While following Ankh amidst his search for his remaining medals, Eiji theorizes as to why Ankh is unable to reform his body. Meanwhile, Kougami discovers Maki's partnership with Kazari and threatens brutal consequences unless Maki ends it. While visiting his sister, Hitomi's grave, Maki calls Kazari to tell him to absorb all of the Core Medals and turn into a Mega Greeed to achieve his desire of ending the world. However, Kazari refuses and creates the Shachi-Panda Yummy from Maki to act on his
Eiji saves Ankh from the foreign Greeed and takes him to Cous Coussier, where they meet with Date to update him on what happened to them. After Date gives Ankh Cell Medals in exchange for the Greeed's origins, Ankh begrudgingly reveals that centuries prior, he helped an ancient king become the original Kamen Rider OOO in his attempts to become a god. Though the king betrayed him to use all of the Greeed's Core Medals to achieve his goal, their power overloaded him and transformed him into the sa