Buffy faces Count Dracula who has come to Sunnydale to make her one of his concubines. Dracula turns Xander into a Renfield of sorts, before being defeated but not killed.
Buffy is having trouble getting along with her sister Dawn, who is kidnapped by Harmony's gang. Harmony makes two inept attempts on Buffy's life, but fails each time and Dawn is saved.
Xander is split into two people by a demon, one strong, the other weak. The lame Xander is unhappy that the cool Xander lives his life better than he can, getting an apartment and generally impressing Anya.
Riley is risking his health to be able to fight with Buffy. His super soldier enhancement is destroying his heart. Buffy enlists Spike to take Riley to the doctor, but instead Spike kidnaps the doctor to remove his chip.
Glory, the new Big Bad, is looking for "The Key". After being beaten by Glory, Buffy meets a monk who reveals to her that she doesn't really have a sister — Dawn is The Key, sent to Buffy for protection.
Tara is about to celebrate her 20th birthday, and her family comes to abduct her before she becomes a demon. Glory sends demons after the Slayer, and Tara accidentally helps them.
After a close call, Buffy asks Spike about the slayers he's killed. Spike tells Buffy the story of his unlife, and reveals a dark secret about Slayers that not even Buffy knew to be true.
Joyce Summers has a brain tumor, and Buffy worries about how Dawn will take it. In addition, Buffy must keep her sister safe from Glory and her magic snake.
As Buffy and Dawn help their mother prepare for surgery, an extraterrestrial preys on Sunnydale's mental patients and follows Joyce home. The alien was summoned by Ben to clean up Glory's mess, and Buffy kills it.
Buffy learns Riley has been going to a vampire brothel to have his blood sucked for pleasure. She confronts him, and he says he's going away with the military if she doesn't give him a reason to stay. She wants him to stay, but doesn't tell him in time.
Emotions between Willow and Anya come to a boil over Xander, and they summon a troll (Olaf). The troll was Anya's ex-boyfriend whom she called vengeance upon. He tries to make Xander choose between the women, but Buffy sends him packing.
The Watcher's Council withholds information about Glory from Buffy and make her go through a series of tests. Glory and the Knights of Byzantium target Buffy, and she realizes it's because she has power over them. She confronts Quentin Travers and the Council members he brought with him, and makes it clear they work for her, not the other way around. Buffy orders Quentin to give her the scoop on Glory, and learns she is not a demon - she's a god.
Dawn discovers that she is the Key, becomes depressed, and runs away. She tells her tale to Ben, who morphs into Glory. Glory, not remembering Ben's thoughts, decides Dawn doesn't know the Key's location, and Willow and Tara teleport Glory elsewhere.
Spike's profession of love for Buffy is spurned when Drusilla returns. He offers to kill Drusilla for her, but Buffy is unimpressed. After Harmony returns to the scene and attacks Spike for revenge, Drusilla tries to kill Buffy. He saves Buffy from Drusilla, but has his invitation to the Summers' home revoked.
A robot girlfriend, April, is abandoned by her creator, Warren. When April tries to kill Warren's current girlfriend, Buffy intervenes. Spike secretly coerces Warren into making him a robot of Buffy.
Buffy and the gang are crushed by the death of Joyce. Dawn goes to the morgue to see the body and is attacked by a vampire. Buffy saves her and watches as Dawn reaches to touch Joyce.
Buffy is comforted by Angel following her mother's death, although he can't stay. Dawn attempts to resurrect Joyce with the help of Spike and Doc, but stops part way through the ceremony after Buffy convinces her.
The Scoobies discover Spike's new robot toy; Glory's minions kidnap Spike, thinking he is the Key, and when they realize he isn't they torture him for information. Buffy and Giles go on a discovery quest about the origin of the Slayer. Buffy is told ominously that "Death is your gift".
Willow and Tara get into an argument, leaving Tara vulnerable to be attacked by Glory, who thinks she is the Key. After discovering her error, Tara's mind is partially sucked out and Willow swears revenge.
Glory discovers the real Key, and Buffy and company must flee from her, and the Knights of Byzantium who are trying to destroy the Key. The gang race from Sunnydale in a caravan.
When Buffy lapses into a catatonic state, Willow uses magic to find out why; a tug-of-war erupts between Glory and Ben over Dawn's ultimate fate.
Buffy and the gang set out to rescue Dawn and fight Glory as the ritual commences. Also, Buffy finally realizes the meaning of "Death Is Your Gift."